Vol.:(0123456789) 1 3
Genetica
DOI 10.1007/s10709-017-9957-y
ORIGINAL PAPER
Genetic diversity and structure of the zombi pea (Vigna vexillata
(L.) A. Rich) gene pool based on SSR marker analysis
Sujinna Dachapak
1
· Prakit Somta
1
· Supalak Poonchaivilaisak
2
· Tarika Yimram
1
·
Peerasak Srinives
1
Received: 9 May 2016 / Accepted: 6 February 2017
© Springer International Publishing Switzerland 2017
America, Africa and Asia, and Australia. NJ tree also sug-
gested that American and Australian accessions are origi-
nated from East African zombi peas, and that the cultivated
accessions from Africa and Asia were genetically distinct,
while those from America were clustered with some cul-
tivated accessions from Africa. These results suggest that
Africa is the center of origin and diversity of zombi pea,
and that domestication of this pea took place more than
once in diferent regions.
Keywords Zombi pea · Diversity · Domestication ·
Microsatellite marker · Underutilized legume
Introduction
The genus Vigna is an agro-sociological important plant
taxon. It comprises more than 100 leguminous plant species
distributed in wide and diverse areas of Africa, America,
Australia and Asia (Lewis et al. 2004). Nine Vigna species
are considered as domesticated crops, including cowpea
(V. unguiculata (L.) Walps.), zombi pea (V. vexillata (L.) A.
Rich), Bambara groundnut (V. subterranean (L.) Verdc.),
mungbean (V. radiata (L.) Wilczek), azuki bean (V. angu-
laris (Ohwi) Ohwi and Ohashi), rice bean (V. umbellata
(Thunb.) Ohwi and Ohashi), black gram (V. mungo (L.)
Hepper), moth bean (V. aconitifolia (Jaqc.) Maréchal),
and créole bean (V. refexo-pilosa Hayata) (Tomooka et al.
2002, 2011). These species are grown mainly for dry seeds
by small farmers in several cropping systems of tropical
and sub-tropical regions. In addition, some other Vigna
species are cultivated and/or harvested for seeds and pods
as food, and for leaves and stems as feedsby local people in
Africa, Asia, Australia and America (Tomooka et al. 2011).
Abstract Zombi pea (Vigna vexillata (L.) A. Rich) is an
underutilized legume species and a useful gene source for
resistance to biotic and abiotic stresses, although there is
little understanding on its genetic diversity and structure.
In this study, 422 (408 wild and 14 cultivated) accessions
of zombi pea from diverse origins (201 from Africa, 126
from America, 85 from Australia, 5 from Asia and 5 from
unknown origin) were analyzed with 20 simple sequence
repeat (SSR) markers to determine its genetic diversity and
genetic structure. The SSR markers detected 273 alleles in
total with a mean of 13.6 alleles per locus. Polymorphism
information content values of the markers varied from
0.58 to 0.90 with an average of 0.76. Overall gene diver-
sity was 0.715. Gene diversity and average allelic richness
was highest in Africa (0.749 and 8.08, respectively) and
lowest in America (0.435 and 4.10, respectively). Nei’s
genetic distance analysis revealed that the highest distance
was between wild Australia and cultivated Africa (0.559),
followed by wild West Africa and wild Australia (0.415).
STRUCTURE, neighbor-joining (NJ), and principal coor-
dinate analyses consistently showed that these zombi pea
accessions were clustered into three major groups, viz.
Electronic supplementary material The online version of this
article (doi:10.1007/s10709-017-9957-y) contains supplementary
material, which is available to authorized users.
* Prakit Somta
pksomta@gmail.com; agrpks@ku.ac.th
1
Department of Agronomy, Faculty of Agriculture
at Kamphaeng Saen, Kasetsart University,
Nakhon Pathom 73140, Thailand
2
Program in Plant Biotechnology, Faculty of Agriculture
at Kamphaeng Saen, Kasetsart University,
Nakhon Pathom 73140, Thailand