Journal of Arid Environments 186 (2021) 104408
0140-1963/© 2020 Elsevier Ltd. All rights reserved.
Plant traits interplay to balance pollen limitation in the Brazilian seasonal
dry forest: A meta-analysis
Thiago Amorim
a
, Andr´ e Maurício Melo Santos
b
, Natan Messias Almeida
c
,
Reinaldo Rodrigo Novo
d
, Ana Virgínia de Lima Leite
d
, Cibele Cardoso Castro
d, e, *
a
Programa de P´ os-Graduaç˜ ao em Ecologia e Recursos Naturais, Departamento de Biologia, Universidade Federal do Cear´ a, Fortaleza, Cear´ a, Brazil
b
Universidade Federal de Pernambuco, Centro Acadˆ emico de Vit´ oria, Vit´ oria de Santo Ant˜ ao, Pernambuco, Brazil
c
Universidade Estadual de Alagoas, Campus de Palmeira dos
´
Indios, Alagoas, Brazil
d
Programa de P´ os-Graduaç˜ ao em Botˆ anica, Departamento de Biologia, Universidade Federal Rural de Pernambuco, Recife, Pernambuco, Brazil
e
Universidade Federal Rural de Pernambuco, Unidade Acadˆ emica de Garanhuns, Garanhuns, Pernambuco, Brazil
A R T I C L E INFO
Keywords:
Caatinga
Floral specialisation
Meta-analysis
Plant habit
Pollen limitation
ABSTRACT
Animal pollination is one of the most important ecosystem services that beneft both natural plant communities
and plant crops. A failure in pollination service can result in pollen limitation of fruit and seed set. This scenario
can be even worse in endangered ecosystems such as tropical dry forests. Pollen limitation is a way of measuring
the reproductive performance of plants by comparing the output of natural pollination with that of supplemental
hand cross-pollination. Studies have shown that pollen limitation can be moderated by plant traits, such as plant
habit, mating system, pollinator dependence and foral specialisation. Herein, we carried out a meta-analysis to
determine the magnitude of pollen limitation in 60 plant species from the Caatinga vegetation of the Brazilian
seasonal dry forest. We found an overall mean pollen limitation of 0.41 and a prevalence of 35%. Trees, shrubs,
and self-incompatible, pollinator dependent and generalist plants were pollen limited. We found a signifcant
interaction between plant habit and foral specialisation. Specialist and generalist trees and generalist shrubs
were pollen limited. Trees and shrubs are mostly threatened in the endangered Caatinga. Species that are not
pollen limited, such as lianas and herbs, may also be indirectly threatened by deforestation and intensive
farming.
1. Introduction
Pollination is a pre-requisite for reproduction in most plants, and
directly impacts their population dynamics, community structure, and
ultimately their ecosystem functioning (Willmer, 2011). Indeed, most
fowering-plants, including crops, need pollinating animals to success-
fully reproduce (Klein et al., 2007; Ollerton et al., 2011). Animal polli-
nation is, thus, an important ecosystem service that is considered in
biological conservation efforts worldwide (Mace et al., 2012). However,
animal pollination is a multifactorial process, involving pre- and
post-pollination events, as well as the environmental stochasticity that
interferes in the population dynamics of pollinators. Consequently,
failure in pollination service can result in inadequate and/or insuffcient
pollen deposition, causing pollen limitation of seed and fruit set (Aizen
and Harder, 2007; Knight et al., 2006). On the other hand, the excessive
deposition of self- or heterospecifc pollen can also have pre- and/or
postzygotic consequences on plant reproductive success (Aizen and
Harder, 2007; Alonso et al., 2013). Although the latter authors argue
that poor-quality receipt plays a more important role than a quantitative
approach, pollen limitation is often assessed by comparing the plant
output (fruit and/or seed set) of natural-pollinated controls with that of
supplemental hand cross-pollinated treatments (Larson and Barrett,
1999; Chen and Zuo, 2018; Sun et al., 2018).
Using this quantitative approach, the magnitude of pollen limitation
has been experimentally determined in plant communities around the
world. Pollen limitation has been found in 25% of the species from a
temperate deciduous forest (Motten, 1986), in 18% of species from a
temperate grassland community (Hegland and Totland, 2008), in 38% of
species from the coastal Atacama desert (Gonz´ alez and P´ erez, 2010), and
in 52% of species from a tropical montane rain forest (Wolowski et al.,
* Corresponding author. Programa de P´ os-Graduaç˜ ao em Botˆ anica, Departamento de Biologia, Universidade Federal Rural de Pernambuco, Recife, Pernambuco,
Brazil.
E-mail address: cibele.castro@ufape.edu.br (C.C. Castro).
Contents lists available at ScienceDirect
Journal of Arid Environments
journal homepage: www.elsevier.com/locate/jaridenv
https://doi.org/10.1016/j.jaridenv.2020.104408
Received 15 April 2020; Received in revised form 25 November 2020; Accepted 27 November 2020