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Resistance of porcine blood clots to lysis relates to
poor activation of porcine plasminogen by tissue
plasminogen activator
Simone M. Flight
a,b
, Paul P. Masci
a,b
, Martin F. Lavin
c
and Patrick J. Gaffney
d
In-vitro experimentation was performed on porcine and
human blood to determine their comparative
responsiveness to a novel fibrinolytic inhibitor and thereby
assess whether the pig is a suitable animal model for
subsequent in-vivo testing of this inhibitor.
Thromboelastography showed the clots formed from
porcine whole blood to be highly resistant to tissue
plasminogen activator (t-PA)-catalyzed lysis, and this
communication offers the resistance of porcine
plasminogen to activation by t-PA as an explanation. Porcine
blood containing 100 and 1500 IU/ml added t-PA lysed very
slowly, having LY30 values of 1.9 W 1.4 and 2.9 W 1.9%,
respectively. In contrast, the LY30 values for the human
clots containing 100 and 1500 IU/ml t-PA were 77.1 W 6.3
and 93.3 W 1.3%, respectively. Moreover, purified porcine
plasminogen was activated very slowly by added t-PA in the
presence of both human and porcine fibrin. Activation of
plasminogen by the endogenous activators, as measured
by the euglobulin clot lysis time, was greatly prolonged for
the pig (22 W 3 h) compared with the human (3.5 W 1.5 h).
These results suggest caution in using the pig as an
experimental model when studying the effects of various
agents on fibrinolysis. Blood Coagul Fibrinolysis 17:417–
420 ß 2006 Lippincott Williams & Wilkins.
Blood Coagulation and Fibrinolysis 2006, 17:417–420
Keywords: fibrinolysis, plasminogen activation, porcine,
thromboelastography
a
School of Medicine, University of Queensland, Princess Alexandra Hospital,
Brisbane,
b
School of Molecular and Microbial Sciences, University of
Queensland, Brisbane,
c
Queensland Institute of Medical Research, Radiation
Biology and Oncology, Royal Brisbane Hospital, Brisbane, Australia and
d
Academic Department of Surgery, St Thomas’ Hospital, London, UK
Correspondence and requests for reprints to Simone Flight, R-Wing, School of
Medicine, Princess Alexandra Hospital, Brisbane, QLD 4102, Australia
Tel: +61 7 3240 2051; fax: +61 7 3240 5063; e-mail: sflight@soms.uq.edu.au
Received 4 January 2006 Accepted 10 March 2006
Introduction
Plasmin is one of the main enzymatic mediators of
fibrinolysis [1,2], catalyzing the cleavage of insoluble
fibrin and thereby accelerating the lysis of a thrombus.
The proenzyme precursor of plasmin is plasminogen
(Plg), a 90-kDa single-chain glycoprotein that is synthe-
sized mainly in the liver and circulates in the plasma at a
concentration of approximately 2 mmol/l [3]. Plasmin
catalyzes the conversion of Glu-Plg to Lys-Plg through
the cleavage of bonds at –K62, –R68 and –K77 [4–7] and
to a greater extent when bound to cell surfaces [8], which
is then converted to two-chain plasmin through the
cleavage of the R561–V562 peptide bond. This conver-
sion to Lys-Plg is a significant but not essential step in the
activation of Glu-Plg in fibrinolysis. The two enzymes
that cleave this bond under physiological conditions are
tissue plasminogen activator (t-PA) and urokinase [9].
While urokinase activation of plasminogen requires no
cofactors, activation with t-PA requires the presence of a
surface and, under physiologic conditions, this surface is
fibrin [10].
Pigs are frequently used to assess the therapeutic effects
of compounds in clinical conditions that involve disrup-
tion of the coagulation, fibrinolysis and the hemodynamic
response. These involve injury [11–13], surgical pro-
cedures [14–16] and chronic disease conditions such as
arteriosclerosis [17]. Accordingly, it is imperative that the
coagulation and fibrinolytic systems of the pig should
approximately mimic those of the human. It has been
reported [18] that pig plasminogen is resistant to acti-
vation by urokinase and that this resistance is related to
differing conformations between human and porcine
plasminogen. Should it be confirmed that bleeding
during surgery is mainly associated with plasmin-
mediated degradation of clots and coagulation factors
[1] then the value of the pig as a model of hemorrhage
would need to be reassessed. This comparative study
investigates the human recombinant t-PA (rt-PA)-stimu-
lated lysis of porcine and human whole blood clots in the
thrombelastograph and compares the relative effects of
endogenous t-PA using the euglobulin clot lysis time
(ECLT) assay for both human and porcine plasmas.
Materials and methods
Materials
rt-PA was obtained from Boehringer Ingelheim (Ingel-
heim, Germany) as Actilyse. Citrated porcine whole
blood and plasma were obtained from the Herston
Medical Research Centre, Brisbane, Australia. Citrated
human blood and plasma was obtained from the Pathol-
ogy unit at Princess Alexandra Hospital, Brisbane.
Lysine-Sepharose affinity resin was purchased from
Amersham Biosciences (Uppsala, Sweden). Bovine
thrombin was purchased from Sigma Aldrich (St Louis,
Missouri, USA).
Short communication 417
0957-5235 ß 2006 Lippincott Williams & Wilkins