Articles www.biosciencemag.org January 2011 / Vol. 61 No. 1฀฀•฀฀BioScience฀฀฀39฀ What Does It Mean to Successfully Conserve a (Vertebrate) Species? KENT H. REDFORD, GEORGE AMATO, JONATHAN BAILLIE, PABLO BELDOMENICO, ELIZABETH L. BENNETT, NANCY CLUM, ROBERT COOK, GUSTAVO FONSECA, SIMON HEDGES, FREDERIC LAUNAY, SUSAN LIEBERMAN, GEOR- GINA M. MACE, AKIRA MURAYAMA, ANDREA PUTNAM, JOHN G. ROBINSON, HOWARD ROSENBAUM, ERIC W. SANDERSON, SIMON N. STUART, PATRICK THOMAS, AND JOHN THORBJARNARSON The conservation of species is one of the foundations of conservation biology. Successful species conservation has often been defined as simply the avoidance of extinction. We argue that this focus, although important, amounts to practicing conservation at the “emergency room door,” and will never be a sufficient approach to conserving species. Instead, we elaborate a positive definition of species conservation on the basis of six attributes and propose a categorization of different states of species conservation using the extent of human management and the degree to which each of the attributes is conserved. These states can be used to develop a taxonomy of species “recovery” that acknowledges there are multiple stable points defined by ecological and social factors. With this approach, we hope to contribute to a new, optimistic conservation biology that is not based on underambitious goals and that seeks to create the conditions under which Earth’s biological systems can thrive. Keywords: species conservation, extinction, successful conservation, recovery Conservation biology as a discipline was not, and is not, the only approach to species conservation. It was grafted to a strong rootstock of species conservation that in some approaches differed from and was overwritten by this emerging discipline. Wildlife management, with its goal of maintaining species in numbers sufficient for significant harvest by humans, has remained largely true to its course and defines successful conservation in terms of harvestable populations. In other approaches, many local and national efforts at species conservation have focused on species with strong human constituencies, irrespective of their conser- vation status. In this article we focus on the interaction between species conservation and conservation biology while recognizing that this framing does not capture the depth or breadth of all species conservation efforts. The conservation biology–based definition of extinction avoidance has become codified in science and policy and has directed conservation largely toward rare and threat- ened species. Extinction avoidance spawned variations of endangered species legislation that focus on what Soulé and colleagues (2003) called “manifest demographic or numeri- cal minimalism.” This perspective is reflected in a statement that attempted to summarize the field of population man- agement: “In conservation, we aim to minimize the chance that a population declines to extinction” (Shea et al. 1998, p. 371). From its origins in population biology and community ecology, conservation biology focused on minimum viable populations and minimum areas necessary to conserve C onservation biology was born as a crisis-oriented and crisis-driven discipline (Meine et al. 2006). Conserva- tion was a natural result of the growing global concern about the loss of tropical forests, coral reefs, and endangered spe- cies, as well as the increasing realization of the truly global impacts of human activities. To gain the attention of the public, decisionmakers, and students, early pioneers of con- servation biology wrote of the sixth extinction, the popu- lation bomb, and the end of nature. Their concerns were contagious, and found fertile soil in a younger generation’s worries about pollution and the decline of iconic species such as whales and giant pandas. Extinction was the middle name of conservation biology, and preventing extinctions was seen as the new discipline’s major aim. The science of modern species conservation came of age with the founding of conservation biology in the 1980s. This foundation was laid in the interplay between genetics and captive breeding (Meine et al. 2006). Among other changes, conservation biology marked a shift in the management of living collections away from displays only and toward population management designed to sustain genetically diverse, demographically stable, and viable captive popula- tions (Hutchins and Smith 2003) that were to serve as assur- ance colonies should wild populations go extinct (Rabb and Saunders 2005). Combined with the crisis discipline per- spective, the three strands of genetics, captive propagation, and crisis were woven into a conservation biology approach to in situ and ex situ species conservation that defined spe- cies conservation as preventing extinction. BioScience 61: 39–48. ISSN 0006-3568, electronic ISSN 1525-3244. © 2011 by American Institute of Biological Sciences. All rights reserved. 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