148 CEREAL CHEMISTRY
Effect of a Cell-Wall-Degrading Enzyme Complex on Starch Recovery
and Steeping Requirements of Sorghum and Maize
Sergio O. Serna-Saldívar
1,2
and Martha Mezo-Villanueva
3
ABSTRACT Cereal Chem. 80(2):148–153
The effect of a commercial cell-wall-degrading enzyme (CWDE)
complex on the steeping time and starch yields of white regular sorghum
(RSOR) compared with yellow maize (YMZ) was determined. An in
vitro wet-milling method standardized to test dosages of 0–120 fungal -
glucanase units (FBG)/100 mL indicated that starch yields were signi-
ficantly higher for YMZ than RSOR and increased proportionally as
enzyme dosage increased. A factorial experiment with a level of con-
fidence of P < 0.05 was performed to study the effect of CWDE addition
to coarsely ground grains for 4 hr after 20 or 44 hr of SO
2
steeping of
whole grains. At both regular steep times, YMZ yielded significantly
higher amounts of starch than RSOR. When steep times were compared,
grains soaked for 48 hr produced 1.7% higher starch yields than counter-
parts treated for 24 hr. CWDE significantly increased starch yields and
recoveries. Enzyme-treated grains yielded 2.5% more starch than
counterparts steeped regularly. For both grains, the best wet-milling
conditions to obtain the highest amount of starch were 48 hr of steeping
and CWDE addition. Under these conditions, YMZ and RSOR yielded
66.9 and 66.6% starch, respectively. Starches obtained after the enzyme
treatment at both steep times contained higher amounts of residual pro-
tein and ash compared with the untreated counterparts. Rapid visco-
amylograph properties of YMZ and RSOR starches were not affected by
the use of the CWDE nor the steep time. In comparison with RSOR
starch, the YMZ starch initiated gelatinization at lower temperature, had
less shear thinning and higher viscosity or setback at the end of cooling.
One of the main industrial uses of maize is for starch pro-
duction. The wet-milling industry has grown because starch is bio-
enzymatically converted into high-fructose corn syrups (HFCS) and
others used in soft drinks. Since the 40s, sorghum has been seen
as a possible alternative to maize for starch (Watson et al 1951,
1955; Watson and Hirata 1955). Sorghum presents several dis-
advantages for wet milling such as the presence of peripheral
endosperm which acts as a barrier against the penetration of the
steep solution, a harder protein matrix and cross-linking which
engulfs starch granules, thus lowering yields and starch quality.
The leaching of phenolic pigments present in the pericarp, testa,
and aleurone tissues produce off-colored starch (Rooney and
Serna Saldivar 2000). Several researchers have found that sorghum
generally yields 10% less prime starch than maize (Watson et al
1951, 1955; Norris and Rooney 1970; Watson 1984; Caransa and
Bakker 1987). Once separated, sorghum starch can be used inter-
changeably with maize starch because both starches have almost
identical viscoamylograph characteristics (Watson 1984; Moheno-
Perez et al 1999).
In Mexico, sorghum is second to maize in terms of area planted
and production, with an estimated production in 2001 of 5.5 million
MT (FAO http//apps.fao.org/). Currently, the relative international
cost of sorghum is 90% the price of maize. However, in Mexico,
the differential has been higher because sorghum is used as animal
feed, whereas maize is channeled directly for human foods. In 2001,
the domestic cost of maize was 20% higher than sorghum (Secre-
taría de Agricultura, Ganadería, Desarrollo Rural, Pesca y Alimen-
tación Centro de Estadística Agropecuaria. www.sagar.gob.mx).
To fulfill demands for maize, during 2001 Mexico imported 5.54
million MT, mainly from the United States, with an estimated value
of 0.5 billion dollars. Therefore, the utilization of sorghum starch
and syrup production can lower maize importation and production
costs. Pioneer research performed by Watson and Hirata (1955)
and Watson et al (1955), in which sorghum was wet-milled in a
manner similar to that of maize, the authors stated that the major
difference is the difficulty of separating sorghum starch and gluten.
In addition, sorghum pericarp is more fragile than the maize
pericarp, thus small pericarp particles impede the clear separation
of the starch and protein and give the starch an off-color. Norris
and Rooney (1970) studied the wet-milling properties of sorghum
and found that the genotype influenced starch recoveries and
quality. Peripheral endosperm content was significantly positively
correlated with protein content of the starch and negatively related
to starch yield and recovery. Steinke and Johnson (1991), Steinke
et al (1991), and Johnston and Singh (2001) demonstrated that the
use of proteolytic and cell-wall-degrading enzymes (CWDE) during
wet-milling of maize significantly lowered SO
2
steeping require-
ments while maintaining similar starch yields. Recently, Johnston
and Singh (2001) effectively used proteases to reduce steep time
and SO
2
requirements in a two-stage wet-milling process. Wang et
al (2000) tested a multiple enzyme cocktail with pectolytic, cellu-
lolytic, hemicellulolytic, and proteolytic activities for wet-milling
of sorghum and found that the enzymes slightly increased starch
yield and produced a more refined starch when compared with
grains steeped with SO
2
. However, the best wet-milling conditions
were achieved with the use of SO
2
and lactic acid.
In a previous experiment, Moheno-Perez et al (1999) found that
maize yielded amounts of starch similar to that of waxy sorghum,
and both grains produced higher amounts of starch than regular or
heterowaxy sorghums. The addition of a fiber-degrading-enzyme
complex to whole grains did not improve the wet-milling process,
with the exception of regular sorghum, where the CWDE reduced
steep time while producing the same starch recovery.
The objectives of this experiment were to compare yields, com-
position, and viscoamylograph properties of starches extracted from
regular sorghum (RSOR) with regular yellow maize (YMZ) and to
determine whether the addition of commercial CWDE to coarsely
ground grains increased starch yields or decreased steep time.
MATERIALS AND METHODS
Grain Utilized
A commercial YMZ with a relatively soft endosperm was used
as the control treatment. The experimental white RSOR (ATX
631*TX436), classed as Type 1, had an intermediate endosperm
texture and regular endosperm. Grains were tested for test weight,
1,000 kernel weight (TKW) and density. Test weight (kg/hL) was
measured with a Winchester bushel meter (Seedburo Equipment,
1
Professor and Department Head, Departamento de Tecnología de Alimentos,
Instituto Tecnológico y de Estudios Superiores de Monterrey, Av. Eugenio Garza
Sada 2501 Sur, CP 64849, Monterrey, N.L. México.
2
Corresponding author. Phone: (81)83284262. Fax: (81)83284322. E-mail:
sserna@itesm.mx.
3
Graduate student, Centro de Biotecnología. Instituto Tecnológico y de Estudios
Superiores de Monterrey, Av. Eugenio Garza Sada 2501 Sur, CP 64849, Mon-
terrey, N.L. México.
Publication no. C-2003-0211-02R.
© 2003 American Association of Cereal Chemists, Inc.