Aquaculture Research. 2020;00:1–8. wileyonlinelibrary.com/journal/are | 1 © 2020 John Wiley & Sons Ltd
1 | INTRODUCTION
Butyl hydroxytoluene (BHT) chemically referred to as 2,6-di-tert-bu-
tyl-p-cresol or 2,6-di-tert-butyl-1-4 methyl phenol is one of the most
extensively used synthetic phenolic antioxidant in foods, mainly con-
taining fats and oils, packaging materials and cosmetics by arresting
the chain reaction in the oxidation of other molecules which results in
increasing shelf-life of the products (Lanigan & Yamarik, 2002; Parke
& Lewis, 1992; Yang et al., 2018). Among various synthetic phenolic
antioxidant, butylated hydroxytoluene (BHT) is most commonly used
in foodstuff with maximum allowable addition amount of 200 mg/Kg
in food, beverages or oil products regulated by the U.S. Food and
Drug Administration (FDA), the European Union (EU) and CODEX
STANDARD (Freitas & Fatibello-Filho, 2010; Yang et al., 2018; Zhou,
Li, Lv, Hu, & Jia, 2015). However, the compound is carried over from
feed to farmed fish (Bohne, Lundebye, & Hamre, 2008). BHT is cur-
rently authorized in feed for all animal species with a maximum con-
centration of 150 mg/kg feed, alone or in combination with another
authorized synthetic antioxidant (Lundebye, Hove, Måge, Bohne, &
Hamre, 2010).
Earlier studies suggested that BHT could exert endo-
crine-disrupting effects, including weak estrogenic activity, the
Received: 23 January 2020
|
Revised: 18 May 2020
|
Accepted: 20 May 2020
DOI: 10.1111/are.14732
ORIGINAL ARTICLE
Toxicity of a synthetic phenolic antioxidant, butyl
hydroxytoluene (BHT), in vertebrate model zebrafish embryo
(Danio rerio )
Raktim Sarmah
1
| Sarada Kanta Bhagabati
1
| Rajdeep Dutta
1
| Dipanka Nath
1
|
Hemanta Pokhrel
1
| Lawonu P. Mudoi
1
| Nishi Sarmah
1
| Jiten Sarma
2
|
Abdul M. Ahmed
1
| Rupam Jyoti Nath
1
| Lucy Ingtipi
3
| Kedolhouse Kuotsu
1
1
Department of Aquatic Environment
Management, College of Fisheries, Assam
Agricultural University, Nagaon, India
2
Department of Fisheries Resource
Management, College of Fisheries, Assam
Agricultural University, Nagaon, India
3
Department of Aquaculture, College of
Fisheries, Assam Agricultural University,
Nagaon, India
Correspondence
Raktim Sarmah, Department of Aquatic
Environment Management, College of
Fisheries, Assam Agricultural University.
Email: rx93@mail.com
Abstract
The synthetic antioxidant 3,5-di-tert-butyl-4-hydroxytoluene (BHT) is widely used as
an additive in the food, cosmetic and plastic industries to increase the tenability of
food and plastic for the past 70 years. BHT is degraded to 3,5-di-test -butyl-4-hydroxy-
benzaldehyde (BHT-CHO) in mammals, as well as in the natural environment such as
in river and water. The average daily intake of BHT for human being is estimated to
be 0.3 mg/kg body weight. Even though it is considered safe for human at authorized
level, but its ubiquitous presence in the aquatic environment and the controversial
toxicological data are of great concern for human as well as aquatic life. The experi-
mental findings of zebrafish embryo toxicity test (ZFET) showed that the acute toxicity
of 96-hr (LC
50
) exposure during the embryogenic stage was found to be 4.388 mg/L
and the effective concentration (EC
50
) was 1.375 mg/L. The reduce heart rate from
the sublethal concentrations indicates the chemical to be cardiotoxic but a further
review is to be needed. The Teratogenic Index (TI) calculated to be 3.19, which implies
the compound may be a potential teratogen in aquatic life. The findings obtained in
this study will stretch more evidence regarding developmental toxicity of BHT, which
will be of much importance in further risk assessment of ecotoxicological studies.
KEYWORDS
BHT, embryotoxicity, teratogenicity, zebrafish