Citation: Alwazeer, D.; Bulut, M.;
Çelebi, Y. Hydrogen-Rich Water Can
Restrict the Formation of Biogenic
Amines in Red Beet Pickles.
Fermentation 2022, 8, 741. https://
doi.org/10.3390/fermentation8120741
Academic Editor: M. Ahsanul Islam
Received: 18 November 2022
Accepted: 9 December 2022
Published: 14 December 2022
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fermentation
Article
Hydrogen-Rich Water Can Restrict the Formation of Biogenic
Amines in Red Beet Pickles
Duried Alwazeer
1,2,3,
* , Menek¸ se Bulut
2,3,4
and Yasemin Çelebi
5
1
Department of Nutrition and Dietetics, Faculty of Health Sciences, Igdir University, Igdir 76000, Turkey
2
Research Center for Redox Applications in Foods (RCRAF), Igdir University, Igdir 76000, Turkey
3
Innovative Food Technologies Development, Application and Research Center, Igdir University,
Igdir 76000, Turkey
4
Department of Food Engineering, Faculty of Engineering, Igdir University, Igdir 76000, Turkey
5
Department of Food Processing, E¸ sme Vocational School, U¸ sak University, U¸ sak 64600, Turkey
* Correspondence: alwazeerd@gmail.com
Abstract: Fermented foods are considered the main sources of biogenic amines (BAs) in the hu-
man diet while lactic acid bacteria (LAB) are the main producers of BAs. Normal water (NW) and
hydrogen-rich water (HRW) were used for preparing red beet pickles, i.e., NWP and HRWP, respec-
tively. The formation of BAs, i.e., aromatic amines (tyramine, 2-phenylethylamine), heterocyclic
amines (histamine, tryptamine), and aliphatic di-amines (putrescine), was analyzed in both beet
slices and brine of NWPs and HRWPs throughout the fermentation stages. Significant differences
in redox value (Eh7) between NWP and HRWP brine samples were noticed during the first and
last fermentation stages with lower values found for HRWPs. Total mesophilic aerobic bacteria
(TMAB), yeast–mold, and LAB counts were higher for HRWPs than NWPs for all fermentation stages.
Throughout fermentation stages, the levels of all BAs were lower in HRWPs than those of NWPs, and
their levels in brines were higher than those of beets. At the end of fermentation, the levels (mg/kg)
of BAs in NWPs and HRWPs were, respectively: tyramine, 72.76 and 61.74 (beet) and 113.49 and
92.67 (brine), 2-phenylethylamine, 48.00 and 40.00 (beet) and 58.01 and 50.19 (brine), histamine, 67.89
and 49.12 (beet) and 91.74 and 70.92 (brine), tryptamine, 93.14 and 77.23 (beet) and 119.00 and 93.11
(brine), putrescine, 81.11 and 63.56 (beet) and 106.75 and 85.93 (brine). Levels of BAs decreased by
(%): 15.15 and 18.35 (tyramine), 16.67 and 13.44 (2-phenylethylamine), 27.65 and 22.7 (histamine),
17.09 and 21.76 (tryptamine), and 21.64 and 19.5 (putrescine) for beet and brine, respectively, when
HRW was used in pickle preparation instead of NW. The results of this study suggest that the best
method for limiting the formation of BAs in pickles is to use HRW in the fermentation phase then replace
the fermentation medium with a new acidified and brined HRW followed by a pasteurization process.
Keywords: red beet (Beta vulgaris) pickle; hydrogen-rich water; molecular hydrogen; biogenic
amines; fermentation
1. Introduction
Red beet is an important source of phytochemicals such as vitamins (C and B), minerals
(K, Fe, P, and Mg), and pigments (betalains) [1]. The pickles of red beets are rich sources
of these bioactive compounds [2]. The fermentation of vegetables in pickle preparation
occurs due to the growth of natural microflora found in raw materials especially Leuconostoc
mesenteroides, Lactobacillus plantarum, Lactobacillus brevis, and Pediococcus [3].
Biogenic amines (BAs) are low molecular weight organic nitrogenous compounds
formed during decarboxylation of amino acids and transamination of aldehydes and
ketones [4]. Most BAs are bioactive, such as tyramine, 2-phenylethylamine, and histamine,
and exert effects on the central nervous system or at a vascular level with pathogenic effects
with different degrees of intensity, ranging from headaches to death. Other amines, such as
putrescine and cadaverine, alter the organoleptic properties of food [5]. Fermented foods
Fermentation 2022, 8, 741. https://doi.org/10.3390/fermentation8120741 https://www.mdpi.com/journal/fermentation