Journal of Phytopathology. 2017;1–16. wileyonlinelibrary.com/journal/jph
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1 © 2017 Blackwell Verlag GmbH
Received: 27 February 2017
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Accepted: 25 April 2017
DOI: 10.1111/jph.12589
ORIGINAL ARTICLE
Potato bacterial wilt suppression and plant health
improvement after application of different antioxidants
Shahenda M. A. Farag
1,2
| Kamel M.A. Elhalag
1,3
| Mohamed H. Hagag
1,2
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Abdel Salam M. Khairy
1,2
| Heba M. Ibrahim
4
| Moheb T. Saker
4
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Nevein A. S. Messiha
1,3
1
Potato brown rot project, Ministry of
Agriculture, Dokki, Giza, Egypt
2
Central Administration of Plant Quarantine,
Dokki, Giza, Egypt
3
Bacterial Diseases Research Department,
Plant pathology research institute, Agricultural
Research Center (ARC), Giza, Egypt
4
Faculty of Agriculture, Physiology
Department, Mansura University, Mansura,
Dakahlia, Egypt
Correspondence
N. A. S. Messiha, Bacterial Diseases Research
Department, Plant Pathology Research
Institute, Agricultural Research Center
(ARC), Giza, Egypt.
Email: nevein_messiha@yahoo.com
Funding information
The Egyptian State Ministry for Scientific
Research, Grant/Award Number: STDF2905
Abstract
Bacterial wilt caused by Ralstonia solanacearum is a devastating disease that often
threatens potato production and exportation. The potential of four antioxidants (sea-
weed extract (SWE), yeast, chitosan and ascorbic acid (ASA)) in controlling the disease
was evaluated in vitro, under glasshouse and field conditions. The field experiment was
conducted in two naturally infested locations: Wardan, Giza (sandy soil), and Talia,
Minufiya (silty clay soil). Only chitosan showed antibacterial properties against the
pathogen in vitro. SWE, yeast and chitosan showed disease suppression under both
glasshouse and field conditions. The disease suppression was accompanied by an in-
crease in the ratio of soil copiotrophic to oligotrophic bacteria. The three antioxidants
increased plant nitrogen content, decreased soil OM content and decreased C/N ratio.
Disease suppression after chitosan application was clearly observed only in Wardan
area, which was characterized by a higher soil alkalinity. A high percentage of antago-
nistic fluorescent strains similar to Pseudomonas putida group were detected for
chitosan-treated plants in Wardan area (sandy soil). ASA drastically decreased the
count of the pathogen in soil, but was conducive to the pathogen in plant tissues. A
remarkable increase in microbial (bacterial and fungal) soil and rhizosphere diversity as
indicated by PCR-DGGE analysis for bacterial 16S rRNA and fungal 18S rRNA was
recorded. In Talia area (silty clay soil), the soil microbial community was more stable
and was in general resistant to the disease where the soils were characterized by high
electrical conductivity. SWE, yeast and ASA significantly increased crop production in
Talia area only.
KEYWORDS
Ralstonia solanacearum, seaweed extract, yeast, chitosan, ascorbic acid, biodiversity, crop
production
1 | INTRODUCTION
Potato is the third most important crop for human consumption after
rice and wheat. Egypt is considered the top potato producer and
exporter in Africa. In 2013, Egypt exported 427,907 ton to Europe,
Russia and Arabic countries as compared to 738,594 ton exported by
whole Africa (FAOSTAT, 2013). Potato brown rot disease (bacterial
wilt) is one of the most important threats for potato production and
international trading. It is a systemic bacterial disease caused by
Ralstonia solanacearum race 3 biovar 2 phylotype II sequevar 1 (Prior
& Fegan, 2005).
In Egypt, potatoes are grown either in the traditional produc-
tion fields in the Nile Delta or in the newly reclaimed areas, mainly
in the desert. Since 1998 and according to FAO standards and EU