VOL. 8, NO. 6, JUNE 2013 ISSN 1990-6145
ARPN Journal of Agricultural and Biological Science
© 2006-2013 Asian Research Publishing Network (ARPN). All rights reserved.
www.arpnjournals.com
457
CHARACTERIZATION AND SELECTION OF HALOPHILIC
MICROORGANISMS ISOLATED FROM MEXICAN SOILS
Delgado-García M.
1
, De la Garza-Rodríguez I.
2
, Cruz Hernández M. A.
3
, Balagurusamy N.
4
, Aguilar C.
1
And Rodríguez-Herrera R.
1
1
Department of Food Research, School of Chemistry, Universidad Autónoma de Coahuila, Blvd. Venustiano Carranza, Saltillo,
Coahuila, Mexico
2
Department of Analytical Chemistry, School of Chemistry, Universidad Autónoma de Coahuila, Blvd. Venustiano Carranza, Saltillo,
Coahuila, Mexico
3
Applied Microbiology Center, GreenCorp Biorganiks de Mexico SA de CV., Saltillo, Coahuila, México
4
School of Biology, Universidad Autónoma de Coahuila, Mexico
E-Mail: raul.rodriguez@uadec.edu.mx
ABSTRACT
Halophilic microorganisms thrive in conditions of high salt concentration and osmotic stress. Due to these
reasons, halophilic microorganisms with ability to fix nitrogen or mobilize phosphate represent a potential as bio-fertilizer
to increase crop growth and yield in saline soils. In this study, 35 different halophilic bacteria were isolated from saline
soils of Coahuila State, Mexico. These bacterial isolates were characterized, and evaluated for their halophilic potential by
growth kinetics, hydrolysis halos formation and in vitro sodium capture. Based on the results, seven potential strains for
saline conditions were selected and were identified by 16S rDNA sequencing analysis Phylogenetic relationship of the
selected strains as well as with other halophilic microorganisms was determined. Six of the identified strains were
Halobacillus trueperi, Bacillus licheniformis, Bacillus pumilus, Staphylococcus succinus, Bacillus atrophaeus and Bacillus
subtilis. Other two strains were identified Halobacillus sp. and Oceanobacillus sp., moderately halophilic organisms.
Keywords: saline soils, 16S rDNA, bacillus spp., halobacillus spp., oceanobacillus sp., staphylococcu succinus.
INTRODUCTION
Worldwide around 897 million of hectares of
surface area is affected by salinity. In Mexico, 10% of the
irrigated area is affected due to high salt concentrations,
and 64% of such saline soils are located in the Northern
region [1]. In soils of arid and semiarid regions, the
problem of excess salts is mainly due to evaporation and
water pressure. In addition, salinity is a problem in areas
with prolonged periods of drought, under dry climate
zones and in coastal areas [2]. This is mainly due to
weathering of mineral and surface of rock crusts, and in
the process get deposited in groundwater resulting in
saline groundwater [3]. Salinity in soils is also due to other
causes such as excessive use of chemical fertilizers, poor
quality of water used for irrigation, deforestation or
intensive cultivation [2]. It is indisputable that high salt
concentrations in soils strongly affect vegetative growth
[4] since high salt concentrations prevent adequate
absorption of nutrients by roots. Besides, high salt content
in soil generates crusts which cause root asphyxia. In
addition, the existence of sodium ion causes dispersion of
organic matter and clays, modifying soil structure [3].
Soil salinity also affects microorganisms.
However, halophilic microorganisms have capacity to
tolerate and live under high salt concentrations by
generating a balance inside and outside of the cell to avoid
osmotic stress [5]. Further, accumulation of compatible
solutes in cytoplasm such as glycine-betaine, sugar
derivate like sucrose or trehalose and/ or inorganic ions
such as sodium, potassium and chloride act as stabilizers
of biologic structures [6, 7]. These traits of halophilic
microorganism generate a great interest to the scientific
community due to their potential applications in the
biotechnological industries [8, 9, 10, 11, 12]. Enzymes of
halophilic microorganisms have advantage over their non-
halophilic counterparts due to their ability to maintain
their stability and catalytic properties at high salt
concentrations [13, 14, 15]. Additionally, the halobacteria
are reported to produce bacteriorrhodopsin, bioplastics,
and various enzymes [16, 17] their potential applications
in different areas.
However, extracellular enzymes production by
halophilic microorganisms has not yet been investigated in
detail. It is generating lot of interest in recent years on
extracellular enzymes like proteases, amylases, DNases,
pullulanases and lipases produced by halophilic
microorganisms [18]. Cuatrociénegas, Coahuila is known
for its high level of soil salinity and presence of abundant
genetic variety of endemic microorganisms. Various
halophilic bacteria have been isolated from small lakes in
Cuatro Ciénegas [19, 20, 21, 22]. However, the halophilic
microorganisms present in the soils of this location and
other areas with saline soils have been little studied [23,
24, 25]. The objective in this study was to isolate and
characterize halophilic microorganisms from saline soils at
different locations of the state of Coahuila, Mexico.
MATERIALS AND METHODS
Soils samples
Twelve soil samples from different saline
locations of Arteaga, Ramos Arizpe, Monclova, Lamadrid
and Cuatro Ciénegas counties of Coahuila, México were
recollected (Table-1). Each sample was taken from the