Vol.:(0123456789) 1 3
Extremophiles
https://doi.org/10.1007/s00792-020-01161-5
ORIGINAL PAPER
Fungi in glacial ice of Antarctica: diversity, distribution
and bioprospecting of bioactive compounds
Graciéle Cunha Alves de Menezes
1
· Bárbara Alves Porto
1
· Soraya Sander Amorim
1
· Carlos Leomar Zani
2
·
Tânia Maria de Almeida Alves
2
· Policarpo Ademar Sales Junior
2
· Silvane Maria Fonseca Murta
2
·
Jeferson Cardia Simões
3
· Betania Barros Cota
2
· Carlos Augusto Rosa
1
· Luiz Henrique Rosa
1
Received: 5 December 2019 / Accepted: 21 February 2020
© Springer Japan KK, part of Springer Nature 2020
Abstract
We identifed cultivable fungi present in the glacial ice fragments collected in nine sites across Antarctica Peninsula and
assessed their abilities to produce bioactive compounds. Three ice fragments with approximately 20 kg were collected, melted
and 3 L fltered through of 0.45 µm sterilized membranes, which were placed on the media Sabouraud agar and minimal
medium incubated at 10 °C. We collected 66 isolates classifed into 27 taxa of 14 genera. Penicillium palitans, Penicillium
sp. 1, Thelebolus balaustiformis, Glaciozyma antarctica, Penicillium sp. 7, Rhodotorula mucilaginosa, and Rhodotorula
dairenensis had the highest frequencies. The diversity and richness of the fungal community were high with moderate
dominance. Penicillium species were present in all samples, with Penicillium chrysogenum showing the broadest distribu-
tion. P. chrysogenum, P. palitans, and Penicillium spp. had trypanocidal, leishmanicidal, and herbicidal activities, with P.
chrysogenum having the broadest and highest capability.
1
H NMR signals revealed the presence of highly functionalized
secondary metabolites in the bioactive extracts. Despite extreme environmental conditions, glacial ice harbours a diverse
fungal community, including species never before recorded in the Arctic and Antarctica. Among them, Penicillium taxa may
represent wild fungal strains with genetic and biochemical pathways that may produce new secondary bioactive metabolites.
Keywords Antarctica · Ice · Fungi · Secondary metabolites
Introduction
Antarctica is one of the most pristine regions in the world
where diferent extreme environmental conditions occur,
which ofer unique opportunities to study the biology of
extremophiles (Rosa et al. 2019). Among the environments
of Antarctica, the Antarctic ice sheet represents the largest
volume of glacial ice on the planet, providing an unknown
ecosystem where biotic factors require special biochemi-
cal mechanisms to survive in low temperatures, low water
activity and nutrient availability conditions, and exposure to
high levels of solar radiation (Anesio and Laybourn-Parry
2012; de Menezes et al. 2019a). The biome of icy environ-
ments seems to be dominated by microorganisms (Sonjak
et al. 2006; Duo Saito et al. 2018; Perini et al. 2019a, b),
which seem to play an important ecological role under the
limit of life (Margesin and Collins 2019). In Antarctica, few
studies have been conducted to recover and identify fungal
species in glacial ice and only a few species of the genera
Alternaria, Aspergillus, Aureobasidium, Candida, Curviba-
sidium, Cystoflobasidium, Davidiella, Penicillium, Crypto-
coccus, Leucosporodium, Pseudozyma, Trichoderma, and
Rhodotorula have been characterized (Jacobs et al. 1964;
Abyzov et al. 2004; D’Elia et al. 2009; Knowlton et al. 2013;
Sanyal et al. 2018). In addition, the glacial Antarctic ice
Communicated by A. Oren.
Electronic supplementary material The online version of this
article (https://doi.org/10.1007/s00792-020-01161-5) contains
supplementary material, which is available to authorized users.
* Luiz Henrique Rosa
lhrosa@icb.ufmg.br
1
Departamento de Microbiologia, Instituto de Ciências
Biológicas, Universidade Federal de Minas Gerais,
Belo Horizonte, MG CEP 31270-901, Brazil
2
Instituto René Rachou, FIOCRUZ-MG, Belo Horizonte, MG,
Brazil
3
Centro Polar e Climático, Universidade Federal do Rio
Grande do Sul, Porto Alegre, Rio Grande do Sul, Brazil