Vol.:(0123456789) 1 3
Mar Biol (2017) 164:79
DOI 10.1007/s00227-017-3114-0
ORIGINAL PAPER
Attachment capacity of the sea urchin Paracentrotus lividus
in a range of seawater velocities in relation to test morphology
and tube foot mechanical properties
Mishal Cohen‑Rengifo
1,2
· Claire Moureaux
2
· Philippe Dubois
2
·
Patrick Flammang
1
Received: 19 October 2016 / Accepted: 17 February 2017
© Springer-Verlag Berlin Heidelberg 2017
calculated a safety factor to predict the fow velocity that
would cause detachment from the substratum. It showed
individuals from the most exposed habitat would resist
higher fow velocities (up to 7.59 ± 0.90 m s
−1
). Both mor-
phometry and tube foot mechanical properties vary among
subpopulations and show an intraspecifc plasticity in P.
lividus. Although, diferences in sea water velocity may
be one cause of this intraspecifc variation, it likely results
from a combination of biotic and abiotic factors.
Abbreviations
σ Stress
Δl Extension
ε Strain
µ Dynamic viscosity
ρ Density
ADV Acoustic Doppler velocimeter
ANOVA Analysis of variance
CC Cap de la Chèvre
C
d
Drag coefcient
C
l
Lift coefcient
F
d
Disk detachment force
F
g
Global detachment force
F
hydro
Hydrodynamic force
F
max
Maximal attachment force
F
t
Traction force
FWL Fractional weight loss
IL Illien
I Length
l
o
Initial length
L Sea urchin length (test diameter with spines)
MO Morgat
PS Polystyrene
PVC Polyvinyl chloride
Re Reynolds number
Abstract Intertidal rocky shores are stressful environ-
ments where benthic invertebrates experience large wave-
induced hydrodynamic forces that can detach them from
the substratum. The tube feet of echinoids counteract these
forces and help them remain securely afxed. Sea urchins
display a high degree of phenotypic plasticity which may
help them cope with hydrodynamic stress. We evaluated
whether habitats presenting diferent seawater velocities
induce plastic responses in the attachment capacity of the
sea urchin Paracentrotus lividus by quantifying their mor-
phology and the adhesive and mechanical properties of
their tube feet. Intertidal adult sea urchins from three sub-
populations were collected around the Crozon peninsula
(France). Localities difered according to measured water
velocities. Size was signifcantly lower in the two most
exposed sites where sea urchins also presented a higher
density of tube feet. Tube foot adhesive properties were
not signifcantly diferent between sites, but their extensi-
bility and toughness were signifcantly higher in individu-
als from the most exposed site. Using this information, we
Responsible Editor: M. Byrne.
Reviewed by undisclosed experts.
* Mishal Cohen-Rengifo
mishal.cohen@student.umons.ac.be
Patrick Flammang
patrick.fammang@umons.ac.be
1
Laboratoire de Biologie des Organismes Marins et
Biomimétisme, Institut des Biosciences, Université de Mons,
23 Place du Parc, 7000 Mons, Belgium
2
Laboratoire de Biologie Marine, (CP160/15), Université
Libre de Bruxelles, 50 Av. F.D. Roosevelt, 1050 Brussels,
Belgium